1- Harvard University (8: Mathematics, Molecular Biology and Genetics, Biology and Biochemistry, Microbiology, Neuroscience and Behavior, Immunology, Psychiatry and Psychology, Clinical Medicine)

2- Massachusetts Institute of Technology (5: Physics, Computer Science, Engineering, Materials Science, Molecular Biology and Genetics)

2- University of California, Berkeley (5: Mathematics, Chemistry, Engineering, Space Science, Environment and Ecology)

4- Kyoto University (Mathematics, Chemistry, Molecular Biology and Biochemistry, Materials Science)

5- Stanford University (3: Mathematics, Computer Science, Engineering)

5- Tokyo University (3: Physics, Chemistry, Biology and Biochemistry)

5- University of California, San Francisco (3: Molecular Biology and Genetics, Biology and Biochemistry, Clinical Medicine)

8- Yale University (2: Neuroscience and Behavior, Psychiatry and Psychology)

8- UCLA (2: Neuroscience and Behavior, Psychiatry and Psychology)

8- University of Wisconsin (Madison) (2: Animal and Plant Science, Agricultural Science)

8- Johns Hopkins University (2: Neuroscience and Behavior, Clinical Medicine)

8- Twin Cities (2: Mathematics, Environment and Ecology)

8- University of Illinois at Urbana-Champaign (2: Computer Science, Engineering)

8- University of California, Davis (2: Zoology, Agricultural Science)